One commonly used classification system for wetlands was developed by Cowardin and is described in Classification of Wetlands and Deepwater Habitats of the United States Exit. The Cowardin system is used by the U.S. Fish and Wildlife Service for the National Wetlands Inventory. In this system, wetlands are classified by landscape position, vegetation cover and hydrologic regime. The Cowardin system includes five major wetland types: marine, tidal, lacustrine, palustrine and riverine.
Another common wetland classification system, used by the Army Corps of Engineers, was developed by Brinson and is described in A Hydrogeomorphic Classification for Wetlands Exit. As the title implies, wetlands are classified by their geomorphic setting, dominant water source (e.g. precipitation, groundwater or surface water) and hydrodynamics. The hydrogeomorphic (HGM) includes five major wetland types: riverine, slope depressional, flat and fringe.
Tidal marsh along the Edisto River, South Carolina.
Common Cattail (Typha latifolia) is a freshwater and estuarine marsh species. Marshes are defined as wetlands frequently or continually inundated with water, characterized by emergent soft-stemmed vegetation adapted to saturated soil conditions. There are many different kinds of marshes, ranging from the prairie potholes to the Everglades, coastal to inland, freshwater to saltwater. All types receive most of their water from surface water, and many marshes are also fed by groundwater. Nutrients are plentiful and the pH is usually neutral leading to an abundance of plant and animal life. We have divided marshes into two primary categories: non-tidal and tidal.
Functions & Values
Marshes recharge groundwater supplies and moderate streamflow by providing water to streams. This is an especially important function during periods of drought. The presence of marshes in a watershed helps to reduce damage caused by floods by slowing and storing flood water. As water moves slowly through a marsh, sediment and other pollutants settle to the substrate or floor of the marsh. Marsh vegetation and microorganisms also use excess nutrients for growth that can otherwise pollute surface water such as nitrogen and phosphorus from fertilizer.
Non-Tidal Marshes
Muskrat (Ondatra zibethicus) next to its house in a cattail-dominated marsh.Description
Non-tidal marshes are the most prevalent and widely distributed wetlands in North America. They are mostly freshwater marshes, although some are brackish or alkaline. They frequently occur along streams in poorly drained depressions and in the shallow water along the boundaries of lakes, ponds and rivers. Water levels in these wetlands generally vary from a few inches to two or three feet, and some marshes, like prairie potholes, may periodically dry out completely.
Highly organic, mineral rich soils of sand, silt, and clay underlie these wetlands, while lily pads, cattails (see photo), reeds and bulrushes provide excellent habitat for waterfowl and other small mammals, such as Red-winged Blackbirds, Great Blue Herons, otters and muskrats. Prairie potholes, playa lakes, vernal pools and wet meadows are all examples of non-tidal marshes.
Functions & Values
Due to their high levels of nutrients, freshwater marshes are one of the most productive ecosystems on earth. They can sustain a vast array of plant communities that in turn support a wide variety of wildlife within this vital wetland ecosystem. As a result, marshes sustain a diversity of life that is disproportionate with their size. In addition to their considerable habitat value, non-tidal marshes serve to mitigate flood damage and filter excess nutrients from surface runoff.
Mink (Mustela vison), a predator of the muskrat.
Pickerelweed (Pontederia cordata).Status
Unfortunately, like many other wetland ecosystems, freshwater marshes have suffered major acreage losses to human development. Some have been degraded by excessive deposits of nutrients and sediment from construction and farming. Severe flooding and nutrient deposition to downstream waters have often followed marsh destruction and degradation. Such environmental problems prove the vital roles these wetlands play. This realization has spurred enhanced protection and restoration of marsh ecosystems, such as the prairie potholes and the Everglades.
Top of Page
Tidal Marshes
The Clapper Rail of the saltmarshes, which is more commonly heard than seen. Description
Tidal marshes can be found along protected coastlines in middle and high latitudes worldwide. They are most prevalent in the United States on the eastern coast from Maine to Florida and continuing on to Louisiana and Texas along the Gulf of Mexico. Some are freshwater marshes, others are brackish (somewhat salty), and still others are saline (salty), but they are all influenced by the motion of ocean tides. Tidal marshes are normally categorized into two distinct zones, the lower or intertidal marsh and the upper or high marsh.
In saline tidal marshes, the lower marsh is normally covered and exposed daily by the tide. It is predominantly covered by the tall form of Smooth Cordgrass (Spartina alterniflora). The saline marsh is covered by water only sporadically and is characterized by Short Smooth Cordgrass, Spike Grass and Saltmeadow Rush (Juncus gerardii). Saline marshes support a highly specialized set of life adapted for saline conditions.
Functions & Values
Tidal marshes serve many important functions. They buffer stormy seas, slow shoreline erosion and are able to absorb excess nutrients before they reach oceans and estuaries. Tidal marshes also provide vital food and habitat for clams, crabs and juvenile fish, as well as offering shelter and nesting sites for several species of migratory waterfowl.
The Great Egret (Casmerodius albus) winters in the tidal marshes along the Gulf Coast.Status
Pressure to fill in these wetlands for coastal development has led to significant and continuing losses of tidal marshes, especially along the Atlantic coast. Pollution, especially near urban areas, also remains a serious threat to these ecosystems. Fortunately, most states have enacted special laws to protect tidal marshes, but diligence is needed to assure that these protective measures are actively enforced.
Top of Page
Skunk Cabbage (Symplocarpus foetidus) sprouts very early in the spring, melting the surrounding snow. The insects that pollinate it are attracted by its odor, which resembles decaying flesh.Description
A swamp is any wetland dominated by woody plants. There are many different kinds of swamps, ranging from the forested Red Maple, (Acer rubrum), swamps of the Northeast to the extensive bottomland hardwood forests found along the sluggish rivers of the Southeast. Swamps are characterized by saturated soils during the growing season and standing water during certain times of the year. The highly organic soils of swamps form a thick, black, nutrient-rich environment for the growth of water-tolerant trees such as Cypress (Taxodium spp.), Atlantic White Cedar (Chamaecyparis thyoides), and Tupelo (Nyssa aquatica). Some swamps are dominated by shrubs, such as Buttonbush or Smooth Alder. Plants, birds, fish, and invertebrates such as freshwater shrimp, crayfish, and clams require the habitats provided by swamps. Many rare species, such as the endangered American Crocodile, depend on these ecosystems as well. Swamps may be divided into two major classes, depending on the type of vegetation present: shrub swamps and forested swamps.
Prothonotary Warblers (Protonotaria citrea) are found in southern swamplands.Functions &Values
Swamps serve vital roles in flood protection and nutrient removal. Floodplain forests are especially high in productivity and species diversity because of the rich deposits of alluvial soil from floods. Many upland creatures depend on the abundance of food found in the lowland swamps, and valuable timber can be sustainably harvested to provide building materials for people.
Status
Due to the nutrient-rich soils present in swamps, many of these fertile woodlands have been drained and cleared for agriculture and other development. Historically, swamps have been portrayed as frightening no-man's-lands. This perception led to the vast devastation of immense tracts of swampland over the past 200 years, such as the destruction of more than half of the legendary Great Dismal Swamp of southeastern Virginia.
Top of Page
Swamps frequently support highly diverse vegetation because of the many layers of vegetation present: shrubs, saplings, and herbaceous plants.Forested Swamps
Forested swamps are found throughout the United States. They are often inundated with floodwater from nearby rivers and streams. Sometimes, they are covered by many feet of very slowly moving or standing water. In very dry years they may represent the only shallow water for miles and their presence is critical to the survival of wetland-dependent species like Wood Ducks (Aix sponsa), River Otters (Lutra canadensis) and Cottonmouth Snakes (Agkistrodon piscivorus). Some of the common species of trees found in these wetlands are Red Maple and Pin Oak (Quercus palustris) in the Northern United States, Overcup Oak (Quercus lyrata) and Cypress in the South, and Willows (Salix spp.) and Western Hemlock (Tsuga sp.) in the Northwest. Bottomland hardwood swamp is a name commonly given to forested swamps in the south central United States.
Top of Page
Button bush (Cephalanthus occidentalis) is found only in shrub swamps.Shrub Swamps
Shrub swamps are similar to forested swamps except that shrubby vegetation such as Buttonbush, Willow, Dogwood (Cornus sp.) and Swamp Rose (Rosa palustris) predominates. In fact, forested and shrub swamps are often found adjacent to one another. The soil is often water logged for much of the year and covered at times by as much as a few feet of water because this type of swamp is found along slow moving streams and in floodplains. Mangrove swamps are a type of shrub swamp dominated by mangroves that covers vast expanses of southern Florida.
Top of Page
This Eastern Mud Salamander (Pseudotriton montanus) is resting on sphagnum moss. Sphagnum creates bogs by holding water and creating acidic conditions. Sphagnum itself may be up to 70 percent water.Description
Bogs are one of North America's most distinctive kinds of wetlands. They are characterized by spongy peat deposits, acidic waters and a floor covered by a thick carpet of sphagnum moss. Bogs receive all or most of their water from precipitation rather than from runoff, groundwater or streams. As a result, bogs are low in the nutrients needed for plant growth, a condition that is enhanced by acid forming peat mosses.
There are two primary ways that a bog can develop: bogs can form as sphagnum moss grows over a lake or pond and slowly fills it (terrestrialization), or bogs can form as sphagnum moss blankets dry land and prevents water from leaving the surface (paludification). Over time, many feet of acidic peat deposits build up in bogs of either origin. The unique and demanding physical and chemical characteristics of bogs result in the presence of plant and animal communities that demonstrate many special adaptations to low nutrient levels, waterlogged conditions, and acidic waters, such as carnivorous plants.
Carlisle Bog in Alaska. Unlike the rest of the United States, Alaska still has most of its wetlands.
Functions and Values
Bogs serve an important ecological function in preventing downstream flooding by absorbing precipitation. Bogs support some of the most interesting plants in the United States (like the carnivorous Sundew) and provide habitat to animals threatened by human encroachment.
Status
Bogs in the United States are mostly found in the glaciated northeast and Great Lakes regions (northern bogs) but also in the southeast (pocosins). Their acreage declined historically as they were drained to be used as cropland and mined for their peat, which was used as a fuel and a soil conditioner. Recently, bogs have been recognized for their role in regulating the global climate by storing large amounts of carbon in peat deposits. Bogs are unique communities that can be destroyed in a matter of days but require hundreds, if not thousands, of years to form naturally.
Nothern Bogs
This bog in Nova Scotia, Canada is dominated by ericaceous dwarf-shrubs, a common family of plants in the peat bogs of the Northeast.
The Northern Pitcher Plant (Sarracenia purpurea) overcomes the nutrient deficiencies of bog life by capturing insects in pools of water in its leaves and digesting them with the help of some local bacteria. The Northern Pitcher Plant's flower looks much like the Sweet Pitcher Plant's (see below).Description
Northern bogs are generally associated with low temperatures and short growing seasons where ample precipitation and high humidity cause excessive moisture to accumulate. Therefore, most bogs in the United States are found in the northern states. Northern bogs often form in old glacial lakes. They may have either considerable amounts of open water surrounded by floating vegetation or vegetation may have completely filled the lake (terrestrialization).
The sphagnum peats of northern bogs cause especially acidic waters. The result is a wetland ecosystem with a very specialized and unique flora and fauna that can grow in these conditions called acidophiles. Nevertheless, bogs support a number of species of plants in addition to the characteristic Sphagnum Moss, including Cotton Grass, Cranberry, Blueberry, Pine, Labrador Tea and Tamarack. Moose, deer, and lynx are a few of the animals that can be found in northern bogs. The Greater Sandhill Crane, the Sora Rail, and the Great Gray Owl depend on bogs for survival.
Top of Page
Pocosins
Pocosins are densely vegetated with trees and shrubs. They are subjected to fire about every 10 to 30 years (Photo by Dr. Curtis Richardson/Duke Wetland Center).
Description
The word pocosin comes from the Algonquin Native American word for "swamp on a hill." These evergreen shrub and tree dominated landscapes are found on the Atlantic Coastal Plain from Virginia to northern Florida; though, most are found in North Carolina. Usually, there is no standing water present in pocosins, but a shallow water table leaves the soil saturated for much of the year.They range in size from less than an acre to several thousand acres located between and isolated from old or existing stream systems in most instances.
Because pocosins are found in broad, flat, upland areas far from large streams, they are ombrotrophic like northern bogs, meaning rain provides most of their water. Also like the bogs of the far north, pocosins are found on waterlogged, nutrient poor and acid soils. The soil itself is a mixture of peat and sand containing large amounts of charcoal from periodic burnings. These natural fires occur because pocosins periodically become very dry in the spring or summer. The fires are ecologically important because they increase the diversity of shrub types in pocosins.
Pocosins provide large tracks of undisturbed land needed by Black Bears (Ursus americanus).
Sweet Bay (Magnolia virginiana.)
Zenobia (Zenobia pulverulenta).The most common plants are evergreen trees (Loblolly Bay, Red Bay and Sweet Bay), and evergreen shrubs (titi, fetterbush and zenobia). Pocosins provide important habitat for many animals, including some endangered species like the red-cockaded woodpecker. They are especially important as the last refuge for Black Bears in coastal Virginia and North Carolina and the Red Wolf has recently been reintroduced in North Carolina pocosins.
Functions and values
Habitat is the most valuable function of Pocosins. Some pocosins are very large and difficult to develop, and so they remain largely undisturbed. As a result, they are a haven for species adapted to living in unaltered forests. As more and more land is developed in the Eastern United States, pocosins are becoming ever more valuable refuges for wildlife.
The Sweet Pitcher Plant (Sarracenia rubra) is one of the carnivorous plants found in pocosins.
The slow movement of water through the dense organic matter in pocosins removes excess nutrients deposited by rainwater. The same organic matter also acidifies the water. This very pure water is slowly released to estuaries, where it helps to maintain the proper salinity, nutrients and acidity. This process is important to help maintain healthy fish populations important to both commerce and recreation. Pocosins are also sources of valuable timber and fuel, but these uses can harm or destroy pocosins if they are not carried out responsibly.
Status
Historically, pocosins were mostly threatened by agriculture. Today, timber harvesting, peat mining, and phosphate mining join agriculture as the biggest threats to the remaining undisturbed pocosins.
Top of Page
Fens, like bogs, are peatlands, but because they are fed by groundwater they are not so acidic as bogs.Description
Fens, are peat-forming wetlands that receive nutrients from sources other than precipitation: usually from upslope sources through drainage from surrounding mineral soils and from groundwater movement. Fens differ from bogs because they are less acidic and have higher nutrient levels. Therefore, they are able to support a much more diverse plant and animal community. These systems are often covered by grasses, sedges, rushes and wildflowers. Some fens are characterized by parallel ridges of vegetation separated by less productive hollows. The ridges of these patterned fens form perpendicular to the downslope direction of water movement. Over time, peat may build up and separate the fen from its groundwater supply. When this happens, the fen receives fewer nutrients and may become a bog.
Showy Lady Slipper, Cypripedium reginae, is an example of a unique plant that thrives in fens.Like bogs, fens are mostly a northern hemisphere phenomenon -- occurring in the northeastern United States, the Great Lakes region, the Rocky Mountains and much of Canada -- and are generally associated with low temperatures and short growing seasons, where ample precipitation and high humidity cause excessive moisture to accumulate.
Functions & Values
Fens, like bogs, provide important benefits in a watershed, including preventing or reducing the risk of floods, improving water quality and providing habitat for unique plant and animal communities.
Fens may be dominated by woody or herbaceous vegetation. In this photograph, trees are invading an herbaceous fen.Status
Like most peatlands, fens experienced a decline in acreage at a rate of about eight percent from 1950 to 1970, mostly from mining and draining for cropland, fuel and fertilizer. Because of the large historical loss of this ecosystem type, remaining fens are that much more rare, and it is crucial to protect them. It is important to recognize that while mining and draining these ecosystems provide resources for people, up to 10,000 years are required to form a fen naturally.
